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    Characters
         separating Hapaline from other palaeotropical 'colocasioid'
      genera  
      (after Grayum, 1990). Comments in [ ] are additional to Grayum‰s
   original list.  
   1. Non-peltate leaves
   (relatively rare in palaeotropical genera) [but cf. Alocasia]  
   2. Spathe-spadix fusion
   [cf. Alocasia aequiloba Engl.] 3. Uniovular ovaries.  
   4. Base chromosome number
   of x = 13 (x = 14 in rest of Caladieae).  
   5. Trinucelate pollen
   (but also present in Remusatia Schott). 
    6. Sterile male flowers
      above and below the fertile male flowers [cf. Alocasia, many species
   with staminodial appendages]  
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